年轻鲍比锹谁拥有焦虑症发现他的治疗狗奥兹实际上是600年的历史，并曾一度由古代强大的巫师所拥有
　　次。在他十二岁生日鲍比被推进到一个新的生命，当他得知他的忠实的宠物真相谁很快决定了男孩和他丧偶
　　母亲的命运。

　　故事发生在1977年的英国伦敦，一位名为谢波德（玛吉·史密斯 Maggie Smith 饰）的老太太开着她的姥爷车来到了一个位于这里的中产社区中。谢波德没有家，吃喝拉撒都在自己的车里，换句话说，在别人的眼中，她就是一名脏兮兮臭烘烘的流浪汉。
　　在这个社区里生活的中产们凭借着他们出色的教养容忍了谢波德女士的入侵，然而他们的善良和宽容却并没有换来谢波德太太的好脸色。这个脾气又倔又犟的老太太让生活在这里的人吃尽了苦头。久而久之，没有人再搭理谢波德太太了，除了一个看上去憨憨的剧作家阿兰（阿历克斯·杰宁斯 Alex Jennings 饰），他不仅每天都对谢波德太太笑脸相迎，甚至同意她占用自己家的车道停车。

　　After breaking out of a rehabilitation zoo, Frankie, a squirrel with all the angles, returns to his forest. He learns from his dim-witted best friend, Cody, a porcupine and street-con, that while he was in lock-up, the entire forest was picked clean by The A.C.O.R.N.S. Institute (Alternative Cosmetic Oils for Regenerative Natural Skin-care). No acorn was left behind. With winter just around the corner, it's a potentially devastating blow to all forest critters. So, Frankie has no choice but to bust into the brand new, state-of-the-art, rodent-proof A.C.O.R.N.S. Processing Plant and take back all of the acorns! Though a seemingly impossible heist, success would not only save the lives of his forest friends, but it would also give Frankie a chance to win back the love of his life, Lola, a local "show squirrel." Sadly, Lola has been burnt by too many of Frankie's false promises in the past. With the support of Frankie's old mentor, Mr. Bellwood, a wealthy badger, Frankie and Cody assemble a team of top notch "crackers" from all over the forest, including: a bat named Fly Boy, a top notch munitions expert; Raitch, a computer hacking albino snake, who never allows her lack of appendages to get in the way; Edsy a tiny "family frog," whose ability to freeze himself solid for days on end without harm is invaluable; a germophobic French rat named Liam, who can collapse his skeleton to get into any space; Chinese father-daughter pangolins named Pango and Lin, the best diggers in the forest; and Bellwood's hedgehog butler, Oscar who knows how to get his hands on anything the crew needs. With winter knocking at their door, Frankie and his crew have only three days to prepare and execute this unprecedented heist. When they learn that the processing of their forest's acorns has been pushed up a day, Lola's refusal to participate leaves Frankie and his crew one critter short. With everything on the line, Frankie and his crew of Animal Crackerz set out to pull off the biggest heist in animal history!

　　布拉德（威尔·法瑞尔 Will Ferrell 饰）是一位电台经理，在邂逅了名为萨拉（琳达·卡德里尼 Linda Cardellini 饰）的女子并同她坠入情网之后，他有了两个可爱的继子梅根（斯嘉丽·埃思特维兹 Scarlett Estevez 饰）和迪伦（欧文·瓦卡罗 Owen Vaccaro 饰），从此，布拉德的人生备忘录上又有了新的的目标——成为两个孩子眼中最棒的继父。
　　然而，一个男人的存在令这个目标看上去遥不可及，那个男人就是萨拉的前夫，孩子们的亲生父亲马龙（马克·沃尔伯格 Mark Wahlberg 饰）。这个看上去不修边幅，整日里游手好闲的男人屡屡骚扰着布拉德的生活。虽然马龙看上去是这么的不靠谱，但他的出现却总是能够受到孩子们的热烈欢迎。